Thanks again gr5. I can now see I have been asking for miracles with my selection of layer height, speed and temperature. I shall now go away and try again while taking account of your recommended throughput/temperature combinations.
I used the Cura slice viewer for the first time and can now see the value in that. The outside shell is being laid down in an anti-clockwise progression. The under-extrusion is therefore happening at the start of a circuit, and this now makes sense to me.