Your extrusion test looks okay indeed, not perfect but should be enough to get good quality at a decent speed.
Regarding your print, i would follow Solid Print 3d's advises, look on the layer view if you can spot something odd. And make sure your model is a good solid (netfabb basic will help checking this).
If everything looks good then there's a problem with either the temperature (too low) or the speed (to high) or both.
If you cleaned the nozzle with the atomic method we can rule that out.
How is your feeder doing? Do you hear some clicking? Does it miss some steps?
One more thing, for such a complex model, wouldn't it be better to cut it in half and print without overhangs? The glue back the two pieces. You would gain in speed and quality.
Assuming you've read all the other topics on feeders, Teflon insulator, general settings, etc....
Have you gone into layer view and looked at what the printer is doing in each layer? Some files I have like that bone you are trying to print are really messed up on the inside, and the printer does some really weird stuff. This has to do with some software exporting the .stl or .obj with voids on the interior. You can go into expert settings and try the different "fix" options on the bottom right of the expert menu and look at the layers, and see how they're printing.
Also, looking at your picture, you're first layer looks terrible! And those layers look like they're .35mm or something! EDIT--> Wow, that came off sounding like a douche, no insult intended
If I were in your shoes, and had to print that model, I would print it in .14mm~.22mm layers, I'd make sure my first layer was perfect. I'd make sure my layer view in Cura was showing a nice clean build. I'd probably open the model in Netfab and do a repair on it. I'd make sure all my settings were good, such as temps and speeds. I'd make sure my retraction settings were dialed in (usually 4.5mm ~ 5mm) by doing a retraction intensive test print (not the cylinder extrusion test).
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